Invasiv art i systemet
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VAD ÄR DETTA?
I den här simuleringen ska du undersöka hur rovdjur (fåglar) och bytesdjur (insekter) påverkas när en ny invasiv art (möss) kommer in i ekosystemet (”invasion”). Antalet individer som introduceras, hur mycket de olika organismerna äter kan varieras. På nästa sida finns uppgifterna. Men först lite mer om hur simuleringen fungerar..
HUR SIMULERINGEN FUNGERAR
Fåglar och insekter rör sig slumpmässigt i rutan som motsvarar ett landskap. Varje steg de rör sig kostar energi och de måste äta någon form av mat. Insekterna äter gräs och fåglarna äter insekter och möss (den invasiva arten). Om en individ får slut på energi (innan den hinner äta) dör den och försvinner ur bild. I den här enkla modellen förökar sig en individ genom att dela sig till två, om den fått i sig tillräckligt med mat. De två som bildas delar på den energi som föräldern hade. Gräset växer till med en viss hastighet, ju mörkare grönt, desto mer energi finns att hämta i en viss fläck. Om gräset nyss ätits upp så finns ingen energi att hämta där (rutan blir vit). Den invasiva arten (möss) kommer in i ekosystemet med knappen ”launch an invasion”. Mössen äter också gräs och konkurrerar med insekterna om födan. Mössen förökar sig på samma sätt som insekterna (med tillräcklig energi delar de sig och blir två, och den energi som föräldern hade fördelas jämnt mellan dem två). Så länge du bara använder ”go/pause” för att starta och stoppa simuleringen fortsätter grafen att ritas med tiden som går. Du kan ändra inställningar (gröna rader med blå reglage) och sedan fortsätta samma simulering. Men om du klickar på ”setup” så börjar grafen om.
HUR DU ANVÄNDER DEN
- Justera parameterarna med reglagen (se nedan), eller använd de inställningar som finns redan.
- Klicka på Uppstart/Omstart.
- Klicka på Start.
- Populationsdynamiken i grafen visar hur antalet individer av olika arter ökar och minskar med tiden.
- I gula rutorna visas aktuellt antal av de olika arterna
- Adjust the NUMBER-INVADERS-TO-ADD and AMOUNT-OF-FOOD-INVADERS-EAT and klicka på LAUNCH INVASION.
- View the POPULATIONS plot to watch the populations fluctuate over time.
Initial Settings:
CONSTANT-SIMULATION-LENGTH: When turned "on" the model run will automatically stop at 2500 ticks. When turned "off" the model run will continue without automatically stopping. AMOUNT-OF-GRASSLAND: The percentage of patches in the world & view that produce grass. INITIAL-NUMBER-BUGS: The initial size of the bug population. INITIAL-NUMBER-BIRDS: The initial size of the bird population. AMOUNT-OF-FOOD-BUGS-EAT: The amount of grass consumed at each simulation step by each one of the bugs. MIN-REPRODUCE-ENERGY-BIRDS: The amount of energy required for a bird to accumulate before it automatically produces an offspring.
Other Settings:
GRASS-TO-BURN-DOWN: Sets the % of designated grassland patches that will have all their grass immediately removed when the BURN THE DOWN GRASS button is klicka påed. %-OF-BUGS-TO-REMOVE: Sets the % of existing bug population that will be immediately removed when the REMOVE BUGS button is klicka påed.
NUMBER-INVADERS-TO-ADD: The number of the invasive species introduced when LAUNCH INVASION is klicka påed. AMOUNT-OF-FOOD-INVADERS-EAT: The amount of grass consumed at each simulation step by each one of the invasive species.
THINGS TO NOTICE
Watch as the grass, predators, and bug populations fluctuate. How are increases and decreases in the sizes of each population related?
How does changing the attributes of the bugs (AMOUNT-OF-FOOD-BUGS-EAT) or birds (MIN-REPRODUCE-ENERGY-BUGS) affect the stability and carrying capacity of the ecosystem?
How does introducing an invasive species that has the same diet as the bugs (i.e. grass) affect the stability of the ecosystem? How does setting the AMOUNT-OF-FOOD-INVADERS-EAT to a different value than AMOUNT-OF-FOOD-BUGS-EAT affect the stability of each population in the ecosystem?
THINGS TO TRY
Try adjusting the parameters under various settings. How sensitive is the stability of the model to the particular parameters. Does the parameter setting affect the amount of fluctuations, the average values of each population, or does it lead to the collapse of one of the populations in the ecosystem (death of all the bugs, birds, or invasive species)?
EXTENDING THE MODEL
Additional invasive plant or invasive predator could be added.
The amount of food a predator or a bug eats could be adjusted so that faster food consumption also corresponds to higher metabolism (cost of energy per move per turn), so that there is a competitive disadvantage for rapid food consumption in some environments.
Traits such as amount of food a predator eats, or min-reproduction energy could be inherited in offspring and subject to slight random changes from mutations.
NETLOGO FEATURES
scale-color is used to visualize the amount of food (energy) available for bugs or invaders at each patch. A whitish green (color of 59.9) is used to represent little grass foliage (not much food available at this patch) and green (color of 55) is used to represents lots of foliage (max. food available at this patch). All other values for food plant-energy are linearly scaled between this range of colors.
RELATED MODELS
Look at the Wolf-Sheep model for a basic predator prey model example.
Look at Bug Hunt Consumers for a simpler version of this model that includes no predators or invaders.
CREDITS AND REFERENCES
This model is a modification of the simulation Bug Hunt Predators and Invasive Species - NetLogo model developed within the BEAGLE curriculum (http://ccl.northwestern.edu/rp/beagle/index.shtml)
The modifications was done 2022 by Ammie Berglund at the Swedish Centre for School Biology and Biotechnolgy (Bioresurs: www.bioresurs.uu.se)
HOW TO CITE
If you mention this model or the NetLogo software in a publication, we ask that you include the citations below.
For the model itself: * Berglund, A. (2022). Invasiv art i systemet. Bioresurs. Modified from:
- Novak, M. and Wilensky, U. (2011). NetLogo Bug Hunt Predators and Invasive Species model. http://ccl.northwestern.edu/netlogo/models/BugHuntPredatorsandInvasiveSpecies. Center for Connected Learning and Computer-Based Modeling, Northwestern University, Evanston, IL.
Please cite the NetLogo software as:
- Wilensky, U. (1999). NetLogo. http://ccl.northwestern.edu/netlogo/. Center for Connected Learning and Computer-Based Modeling, Northwestern University, Evanston, IL.
COPYRIGHT AND LICENSE
Copyright 2011 Uri Wilensky.
This work is licensed under the Creative Commons Attribution-NonCommercial-ShareAlike 3.0 License. To view a copy of this license, visit https://creativecommons.org/licenses/by-nc-sa/3.0/ or send a letter to Creative Commons, 559 Nathan Abbott Way, Stanford, California 94305, USA.
Commercial licenses are also available. To inquire about commercial licenses, please contact Uri Wilensky at uri@northwestern.edu.
Comments and Questions
globals [ birds-color ;; color of birds bugs-color ;; color of bugs grass-color ;; color of patches/grass dirt-color ;; color of areas that have no grass right now, but will grow grass back shortly invasive-color ;; color of invaders bugs-stride ;; how much a bug moves in each simulation step invaders-stride ;; how much an invader moves in each simulation step birds-stride ;; how much a bird moves in each simulation step bird-size ;; size of birds bug-size ;; size of bugs invader-size ;; size of invaders bug-reproduce-age ;; min age of bugs before they can reproduce invader-reproduce-age ;; min age of invaders before they can reproduce bird-reproduce-age ;; min age of birds before they can reproduce max-bugs-age ;; max age of bugs before they automatically die max-birds-age ;; max age of birds before they automatically die max-invaders-age ;; max age of invaders before they automatically die birds-energy-gain-from-bugs ;; how much energy birds gain from eating bugs birds-energy-gain-from-invaders ;; how much energy birds gain from eating invaders min-reproduce-energy-bugs ;; energy required for reproduction by bugs min-reproduce-energy-invaders ;; energy required for reproduction by invaders max-bugs-offspring ;; maximum number of offspring per reproductive event for a bug max-birds-offspring ;; maximum number of offspring per reproductive event for a bird max-invaders-offspring ;; maximum number of offspring per reproductive event for an invader max-plant-energy ;; maximum amount of energy a plant can "grow" to store in a patch sprout-delay-time ;; delay time before a plant begins to grow back, after all of its foliage has been eaten plant-growth-rate ;; amount of energy the plant stores as food per tick as it grows back ] breed [ bugs bug ] breed [ invaders invader ] breed [ birds bird ] breed [ disease-markers disease-marker ] ;; turtles that mark the location of where a bug was removed from disease. breed [embers ember] ;; turtles that mark the location of the flames and embers of the fire. turtles-own [ energy current-age max-age ] ;; these three variables are used for bugs, invaders, and birds ;; fertile? are patches that can grow grass ;; plant-energy keeps track of the amount of food or foliage at that patch ;; countdown keeps track of how much longer until the sprout-delay-time has elapsed for this patch patches-own [ fertile? plant-energy countdown] to setup clear-all set max-plant-energy 100 set plant-growth-rate 10 set sprout-delay-time 25 set bugs-stride 0.3 set invaders-stride 0.3 set birds-stride 0.4 ;; a larger birds-stride than bugs-stride leads to scenarios where less birds ;; can keep a larger number of bugs in balance. set bug-reproduce-age 20 set invader-reproduce-age 20 set bird-reproduce-age 40 set max-birds-age 100 set max-invaders-age 100 set max-bugs-age 100 set birds-energy-gain-from-bugs 25 set birds-energy-gain-from-invaders 25 set min-reproduce-energy-bugs 10 set min-reproduce-energy-invaders 10 set max-bugs-offspring 3 set max-invaders-offspring 3 set max-birds-offspring 2 set bird-size 2.5 set bug-size 1.2 set invader-size 1.5 set birds-color (red - 1) set bugs-color (violet) set invasive-color (blue - 3) set grass-color (green) set dirt-color (white) set-default-shape bugs "bug" set-default-shape birds "bird" set-default-shape invaders "mouse" add-starting-grass add-bugs add-birds reset-ticks end to go if ticks >= 1000 and simulera-endast-1000-tidssteg? [ stop ] ask disease-markers [ age-disease-markers ] ask turtles [ ;; we need invaders and bugs to eat at the same time ;; so one breed doesn't get all the tasty grass before ;; the others get a chance at it. (ifelse breed = bugs [ bugs-live reproduce-bugs ] breed = invaders [ invaders-live reproduce-invaders ] breed = birds [ birds-live reproduce-birds ] [ ] ; anyone else doesn't do anything ) ] ask patches [ ;; only the fertile patches can grow grass set countdown random sprout-delay-time grow-grass ] tick end to remove-a-%-bugs let number-bugs count bugs ask n-of floor (number-bugs * %-minskning-insekter / 100) bugs [ hatch-disease-markers 1 [ set current-age 0 set size 1.5 set color red set shape "x" ] die ] end to age-disease-markers set current-age (current-age + 1) set size (1.5 - (1.5 * current-age / 20)) if current-age > 25 or (ticks = 999 and simulera-endast-1000-tidssteg?) [die] end to start-fire ask patches [ set countdown sprout-delay-time set plant-energy 0 color-grass] end to add-starting-grass ;; setup patch procedure to add grass based on initial settings let number-patches-with-grass (floor (andel-markvegetation * (count patches) / 100)) ask patches [ set fertile? false set plant-energy 0 ] ask n-of number-patches-with-grass patches [ set fertile? true set plant-energy max-plant-energy / 2;; (max-plant-energy - random max-plant-energy) ] ask patches [color-grass] end to add-bugs ;; setup bugs procedure to add bugs based on initial settings create-bugs antal-insekter-vid-start [ ;; create the bugs, then initialize their variables set color bugs-color set energy 20 + random 20 - random 20 ;;randomize starting energies set current-age 0 + random max-bugs-age ;;don't start out with everyone at the same age set max-age max-bugs-age set size bug-size setxy random world-width random world-height ] end to add-birds ;; setup birds procedure create-birds antal-fåglar-vid-start ;; create the bugs, then initialize their variables [ set color birds-color set energy 40 + random 40 - random 40 ;;randomize starting energies set current-age 0 + random max-birds-age ;;don't start out with everyone at the same age set max-age max-birds-age set size bird-size setxy random world-width random world-height ] end to add-invaders create-invaders antal-av-invasiv-art-som-läggs-till ;; create the bugs, then initialize their variables [ set color invasive-color set energy 20 + random 20 - random 20 ;;randomize starting energies set current-age 0 + random max-invaders-age ;;don't start out with everyone at the same age set max-age max-invaders-age set size invader-size setxy random world-width random world-height ] end to bugs-live ;; bugs procedure move-bugs set energy (energy - 1) ;; bugs lose energy as they move set current-age (current-age + 1) bugs-eat-grass death end to birds-live ;; birds procedure move-birds set energy (energy - 1) ;; birds lose energy as they move set current-age (current-age + 1) birds-eat-prey death end to invaders-live ;; invaders procedure move-invaders set energy (energy - 1) ;; invaders lose energy as they move set current-age (current-age + 1) invaders-eat-grass death end to move-bugs ;; bugs procedure rt random 50 - random 50 fd bugs-stride end to move-birds ;; birds procedure rt random 50 - random 50 fd birds-stride end to move-invaders ;; invaders procedure rt random 40 - random 40 fd invaders-stride end to bugs-eat-grass ;; bugs procedure ;; if there is enough grass to eat at this patch, the bugs eat it ;; and then gain energy from it. if plant-energy > hur-effektivt-insekterna-äter [ ;; plants lose ten times as much energy as the bugs gains (trophic level assumption) set plant-energy (plant-energy - (hur-effektivt-insekterna-äter * 10)) set energy (energy + hur-effektivt-insekterna-äter) ;; bugs gain energy by eating ] ;; if plant-energy is negative, make it positive if plant-energy <= hur-effektivt-insekterna-äter [set plant-energy 0] end to invaders-eat-grass ;; invaders procedure ;; if there is enough grass to eat at this patch, the invaders eat it ;; and then gain energy from it. if plant-energy > hur-effektivt-invasiva-arten-äter [ ;; plants lose 10 * as much energy as the invader gains (trophic level assumption) set plant-energy (plant-energy - (hur-effektivt-invasiva-arten-äter * 10)) set energy energy + hur-effektivt-invasiva-arten-äter ;; bugs gain energy by eating ] ;; if plant-energy is negative, make it positive if plant-energy <= hur-effektivt-invasiva-arten-äter [set plant-energy 0] end to birds-eat-prey ;; birds procedure if (any? bugs-here and not any? invaders-here) [ask one-of bugs-here [die] set energy (energy + birds-energy-gain-from-bugs) ] if (any? invaders-here and not any? bugs-here) [ask one-of invaders-here [die] set energy (energy + birds-energy-gain-from-invaders) ] if (any? invaders-here and any? bugs-here) [ifelse random 2 = 0 ;; 50/50 chance to eat an invader or a bug if both are on this patch [ask one-of invaders-here [die] set energy (energy + birds-energy-gain-from-invaders) ] [ask one-of bugs-here [die] set energy (energy + birds-energy-gain-from-bugs) ] ] end to reproduce-bugs ;; bugs procedure let number-offspring (random (max-bugs-offspring + 1)) ;; set number of potential offspring from 1 to (max-bugs-offspring) if (energy > ((number-offspring + 1) * min-reproduce-energy-bugs) and current-age > bug-reproduce-age) [ if random 2 = 1 ;;only half of the fertile bugs reproduce (gender) [ set energy (energy - (number-offspring * min-reproduce-energy-invaders)) ;;lose energy when reproducing- given to children hatch number-offspring [ set size bug-size set color bugs-color set energy min-reproduce-energy-bugs ;; split remaining half of energy amongst litter set current-age 0 rt random 360 fd bugs-stride ] ;; hatch an offspring set it heading off in a random direction and move it forward a step ] ] end to reproduce-invaders ;; bugs procedure let number-offspring (random (max-invaders-offspring + 1)) ;; set number of potential offspring from 1 to (max-invaders-offspring) if (energy > ((number-offspring + 1) * min-reproduce-energy-invaders) and current-age > invader-reproduce-age) [ if random 2 = 1 ;;only half of the fertile invaders reproduce (gender) [ set energy (energy - ( number-offspring * min-reproduce-energy-invaders)) ;;lose energy when reproducing - given to children hatch number-offspring [ set size invader-size set color invasive-color set energy min-reproduce-energy-invaders ;; split remaining half of energy amongst litter set current-age 0 rt random 360 fd invaders-stride ] ;; hatch an offspring set it heading off in a random direction and move it forward a step ] ] end to reproduce-birds ;; bugs procedure let number-offspring (random (max-birds-offspring + 1)) ;; set number of potential offspring from 1 to (max-invaders-offspring) if (energy > ((number-offspring + 1) * energinivå-för-fortplantning-fågel) and current-age > bird-reproduce-age) [ if random 2 = 1 ;;only half of the fertile bugs reproduce (gender) [ set energy (energy - (number-offspring * energinivå-för-fortplantning-fågel)) ;;lose energy when reproducing - given to children hatch number-offspring [ set current-age 0 set size bird-size set color birds-color set energy energinivå-för-fortplantning-fågel;; split remaining half of energy amongst litter set current-age 0 rt random 360 fd birds-stride ] ;; hatch an offspring set it heading off in a random direction and move it forward a step ] ] end to death ;; common procedure for bugs, birds & invaders to die ;; die when energy dips below zero (starvation), or get too old if (current-age > max-age) or (energy < 0) [ die ] end to grow-grass ;; patch procedure set countdown (countdown - 1) ;; fertile patches gain 1 energy unit per turn, up to a maximum max-plant-energy threshold if fertile? and countdown <= 0 [set plant-energy (plant-energy + plant-growth-rate) if plant-energy > max-plant-energy [set plant-energy max-plant-energy]] if not fertile? [set plant-energy 0] if plant-energy < 0 [set plant-energy 0 set countdown sprout-delay-time] color-grass end to color-grass ;; patch procedure ifelse fertile? [ ifelse plant-energy > 0 ;; scale color of patch from whitish green for low energy (less foliage) to green - high energy (lots of foliage) [set pcolor (scale-color green plant-energy (max-plant-energy * 2) 0)] [set pcolor dirt-color] ] [set pcolor dirt-color] end ; Copyright 2011 Uri Wilensky. ; See Info tab for full copyright and license.
There is only one version of this model, created over 3 years ago by Ammie Berglund.
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